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TCP reliable delivery depends on two fields in the TCP header: the sequence number and the acknowledgement number. The sequence number identifies the first byte of data in each segment, and because every byte in a transmission is numbered sequentially from a randomly chosen starting point, both sides of the connection can track exactly which data has been received and what comes next. When large payloads are broken into multiple segments at Layer 4, each segment carries the sequence number of its first byte, giving the receiving device everything it needs to reassemble the full data stream in the correct order even if segments arrive out of sequence.
The acknowledgement mechanism works alongside sequencing to confirm successful delivery and trigger retransmission when needed. After receiving one or more segments, the destination device sends back an acknowledgement number representing the next byte it expects, effectively confirming receipt of all data up to that point. The sender reads this acknowledgement, confirms which segments got through, and continues transmitting from the appropriate sequence number. If the receiver detects a gap, because an expected sequence number never arrives, it sends an acknowledgement for the last successfully received byte, prompting the sender to retransmit the missing segment.
Both sides of a TCP connection maintain their own independent sequence and acknowledgement numbers, since data typically flows in both directions. Tools like Wireshark may normalize sequence numbers to zero for readability, but the actual values in the header are large numbers drawn from a 32-bit field, allowing over four billion possible byte labels before the sequence space wraps around. This combination of sequencing, acknowledgement, and retransmission is what makes TCP a reliable transport protocol, capable of delivering complete, ordered data even across unreliable network paths.
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